Do they feel hollow or thin around the chest, are they loosing weight or condition?
How old are the birds?
Where did they come from? Could you ask if the breeder has any others with this problem at the moment?
Additional vitamins may help. There is a children's vitamin product called Abidec, a few squirts in the water might help.
I had a similar problem with Barnvelders years ago and lost a breeding trio from a local breeder. They had the same problem as yours but with the additional symptoms above.
I hope they are perking up for you, you seem to be giving them the best care they could have.
Don't blame yourself, sometimes rare breeds and fancy hens are a bit deficient with immunity due to close breeding and matching.
Another thought.... Do you have good ventilation in the coop? They need a bit of air flow.